The Bruins had dropped three straight to the Trojans under Jim Mora Jr before he was sacked on Sunday Nov. 19.
"Making a coaching change is never easy", UCLA athletic director Dan Guerrero said in a statement, "but it's an especially hard decision when you know that a coach has given his all to our university".
Athletic director Dan Guerrero announced the shocking move one day after rival Southern California's 28-23 victory over UCLA (5-6, 3-5 Pac-12). "We thank Jim and his family for his service to our school and his unquestionable commitment to our student-athletes".
In what would become just short of a six-year stint at UCLA, Mora produced an overall record of 46-30.
Every UCLA Head Football Coach has managed a double-digit winning season, besides Rick Neuheisel, since Terry Donahue took over in 1976 but the Bruins haven't been ranked number 1 since 1954.
The Bruins' statement noted that Mora's contract will be bought out exclusively using department-generated funds. Given that UCLA had only one top-25 finish between 1999 and 2012, Mora appeared to have the Bruins on the right track, especially with the arrival of five-star quarterback prospect Josh Rosen to replace Brett Hundley in 2015. The Bruins also announced that they would conduct a "national search" for the next head coach, with a group that includes Troy Aikman participating in finding the next leader of the program. UCLA missed the postseason previous year after a 4-8 finish.
Offensive coordinator Jedd Fisch has been installed as interim coach.
UCLA coach Jim Mora looks on as his Bruin team is defeated by USC on November 18, 2017 at the Coliseum. Through 11 games, UCLA ranks fourth in the Pac-12 in points per game (34.1) and third in total offense (466.0 yards per game).